Kyle Szczepaniuk (author), Michigan State University, James Harding (editor), Michigan State University, Rachelle Sterling (editor), Special Projects. Relative to its size, the jaguar has the most powerful bite of big cats, registering at 1,500 PSI. Nile monitors were added to Floridas Prohibited species list. Given that the average Nile croc grows to 16 feet, it stands to reason that their actual bite strength should measure upwards of 3,000 PSI a value which exceeds that of the American alligator, bite-force wise. Hekkala. Lateral view of dentition of adult Nile monitor (Varanus niloticus), showing rounded teeth at the back of the jaws. 2000. The University of Chicago's Porro noted that no Tyrannosaurus rex muscle survives, so estimates for the dinosaur's bite force are based on its body size, wide skull 6.63) is a large species and reaches a length of about 1.5m. There are 89 premaxillary teeth, 1213 teeth on each maxilla, and 1113 teeth on each dentary. All these pig species pile up a nest when due to give birth. It has a varied diet including arthropods, mollusks, eggs, birds, and snakes. A very spectacular group of birds, the birds's of paradise (Fam. Thank you for reading! (Branch, 1998; Cowles, 1928; Steel, 1996), Estimates vary, but the expected lifespan in captivity is reported at between 10 and 20 years. Due to their generalist diet, Nile monitors may impact state- and federally-listed species including sea turtles, wading birds, gopher tortoises and the American crocodile. The female will excavate a secure place to lay her clutch, either in the ground or in an active termite nest. Varanid skulls are mesokinetic, possibly amphikinetic, and the snout can be raised or lowered relative to the cranium. In areas where Nile monitors are abundant, they have also attacked small pets and livestock such as chickens. Mertens, R. (1942). Report injured, orphaned or dead manatees, Florida Youth Conservation Centers Network, Conditional & Prohibited Nonnative Species Regulation, Regulations for Conditional Snakes and Lizards, Nonnative Species Permit Applications and Information, See a full list of our Social Media accounts. The other clade includes the monitors of the Sahel (Mali to Ethiopia) and Nile regions. Therefore, a patient who is at risk for eosinophilic meningitis from A. cantonensis should have blood tests to measure the level of eosinophil leukocytes before performing a lumbar puncture. On the subcontinent , they occur through the eastern part of the region, extending along the Orange River to the Atlantic Ocean and Fish River Canyon, and along the South coast to Gamtoos River Valley, but is absent from the Western Cape. The larger V. mertensi is confined to freshwater and shows significant adaptations to the aquatic habitat: it has pronounced lateral compression of the elongated tail, nostrils situated on top of the snout that become sealed when the animal submerges, and an ability to remain active at low body temperatures (Bennett, 1995). Its distribution range is limited to northern Australia while V. indicus is more widespread occurring in New Guinea and parts of Southeast Asia. Some monitors, such as the African monitor (Varanus niloticus) spend a considerable amount of time in and around shallow water (Lenz, 1995; de Buffrnil and Hmery, 2002). Occlusal views of the upper and lower dentitions of Nile monitor (Varanus niloticus), showing the conical grasping teeth at the front of the jaws and rounded crushing teeth at the back. Monitor lizards Varanus salvator occur in very different ecosystems, they are excellent swimmers and juveniles and semiadults are also good climbers. Radiological examination is not specific for angiostrongyliasis. Are Nile Monitors Dangerous Although monitor lizards are capable of producing venom, there is no danger of them dying in the wild. This is primarily due to their extremely territorial and aggressive nature. They are not found in any of the desert regions of Africa, however, as they thrive around rivers. In the absence of competition, the Komodo dragon is an aggressive and fearless hunter and the adults prey on large mammals, such as deer. Even humans can bite with a great deal of force. Neighbour JeffWildonger, now aged 38, told The Sun Online: "If he didn't have those lizards there he'd be alive today, there's no doubt.". Antigen with a molecular weight of 29 or 31kDa is highly specific for A. cantonensis (Maleewong et al., 2001; Eamsobhana and Yong, 2009). Monitors, Tegus, and Related Lizards. Nile monitor lizards consume crocodile eggs and may act as population control in some areas. This material is based upon work supported by the He described it again in 1766 as Lacerta nilotica. In parts of Africa and Asia, varanids are hunted for their skins or as medicinal ingredients, but mainly for their meat. Humans may not be able to compete one-on-one with any of these powerful predators but we are still their greatest threat. Mating and egg laying usually follows the rainy season, which varies regionally. [2] The long-tailed macaques (Macaca fascicularis) occur in closed forests as well as in mangroves and feed on fruits, leaves, bark, arthropods (including crabs) and molluscs. They may occasionally work in pairs to rob a crocodile nest, with one lizard distracting the guarding mother crocodile while the other races in to grab eggs. This species is listed under CITES, Appendix II. National Science Foundation A detailed account on the turtle nesting sites in the Andaman and Nicobar Islands was reported by Andrews and Tripathy (2004). Here, semi-aquatic varanids include the Large water monitor (V. mertensi), and the Mangrove monitor (V. indicus). "Varanus niloticus" (On-line), Animal Diversity Web. The majority of these lizards follow a carnivorous diet and feed off the local wildlife in their range. This action drags the teeth through the flesh, distal side first. How frequently do Nile monitors eat? Not every monitor lizard is giant, but some are capable of competing with the largest lizards in the world in size. Lets find out how dangerous these giant reptiles are. Teeth are absent from the palate (Mahler and Kearney, 2006). San Diego, California: Academic Press. The skull of the Komodo dragon seems to be adapted to minimizing stresses set up by pulling forces on the jaws rather than to exerting a large bite force (Moreno etal., 2008; Fry etal., 2009). The Nile crocodile is considered the deadliest crocodile species in the world due to its territorial and aggressive nature. Generally, grizzlies live for about 25 years in the wild, and they grow to between five and eight feet tall while tipping the scales at almost 800 pounds. Instead of dragging down and eviscerating their lunch, or biting the throat like most other large felines, the jaguar crushes the skull of their victims by biting right through it. Nile monitors are native to Africa and the species is distributed throughout the entire central and southern regions of the continent, including Sudan and a portion of central Egypt along the Nile River. As a last resort a cornered Nile monitor will bite savagely and eject foul-smelling matter from the cloaca to deter attack or throw off a pursuer. & De Villiers, M.S. These brilliantly ornamented males engage themselves in sometimes very elaborate ritual dances on special trees or dancing grounds to attract a mate. They are arboreal and use the strong grip of their mouth to hold onto prey while climbing trees. This season correlates with the development of the gonads. Buffrenil, V., G. Hemery. 2002. In monitor lizards that include a large proportion of shelled prey in their diet as adults, the posterior teeth of adults may be wider and blunter than those in juveniles, an adaptation to durophagy. Buffrenil (1992) considered that, when fighting for its life, a Nile monitor was a more dangerous adversary than a crocodile of a similar size. Nile monitors may escape confinement by pushing off the tops of cages or by using their sharp claws to tear through screens. Depending on size, the diet can include mammals, birds, reptiles, fishes, arthropods, eggs, crustaceans, and mollusks. But a close second is the tiger, the largest wild cat in the world, with a bite force of 1,050 PSI. Large volume and highly compliant lungs give an increased tidal volume and low respiratory rate (Bellairs 1969d; Bennett, AF 1972; Perry 1989; Perry & Duncker 1978). (Anadu, 1987; Bartlett and Bartlett, 1996; Branch, 1998), Nile monitors are sometimes known as "water monitors", or "water leguaan" in South Africa. body mass of 4.7kg (10lb). In other parts of the world, the reptiles known as marine iguanas were found, whereas it is are not found in Andaman and Nicobar Islands. Lystrosaurus was one of the few terrestial species that survived the permian extinction. The salt water crocodile (Crocodylum porosus) and water monitor lizard (Varanus salvator) are found living in the mangrove areas of Andaman and Nicobar Islands. Just check out this article about the most venomous mammals in the world. The marine turtlesfound in Andaman and Nicobar Islands consist of four species namely Leatherback turtle (Demochelys coriacea), Green turtle (Chelonia mydas), Hawsbill turtle (Eretmocheylys imbricata squamota), and Olive Ridley turtle (Lepidochelys olivacea). They engage in mutual tongue-licking and grooming, dewlap extension and body rubbing, and at night they often sleep together on branches. Bates, M.F., Branch, W.R., Bauer, A.M., Burger, M., Marais, J., Alexander, G.J. With an average length of 100-140 cm (around 4 feet) and a maximum of 200 cm (6 1/2 feet), it is the largest African lizard. The Nile monitors high reproduction rate, diverse diet, and ability to travel over land and in fresh and saltwater allows for potential establishment throughout Florida, particularly in coastal areas dominated by mangroves or salt marshes. By continuing you agree to the use of cookies. For further details of our complaints policy and to make a complaint please click this link: thesun.co.uk/editorial-complaints/, Ronald Huff, 42, cuddles one of his pet monitor lizards, which then ate him, It later emerged the lizards had been allowed to roam freely throughout his home, The former soldier's facewas eaten by seven of the 6ft-long monsters, Ronald's pal Jeff Wildonger drew this sketch of the scene the night after he saw the half-eaten corpse of his neighbour, Ronald's former boss Mike also claimed the huge creatures had attacked Ronald in the past, Independent Press Standards Organisation (IPSO). Postal: Private Bag X101, Silverton, 0184. When walking and foraging, the Nile monitor holds its body well off the ground, and uses its long, forked tongue to sense potential food sources. 9 Incredible Lizards That Look Like Dragons, Monitor Lizard vs Komodo Dragon: Whats the, The 5 Best Vitamin Supplements for Geckos. Crocodiles also take monitors on occasion. This family is comprised of one genus (Varanus) and nearly 70 species of monitor lizard. It has a typical varanid dentition, with sharp, pointed, spaced teeth (Fig. Some animals can not be categorized in a special feeding guild as herbivore, insectivore or carnivore, since they forage on food belonging to all or a combination of these categories. The ADW Team gratefully acknowledges their support. View our online Press Pack. They found to migrate long distances in the sea and reach sandy coastal areas to lay their eggs. On this Wikipedia the language links are at the top of the page across from the article title. 2016. 4.20). [12] In 2016, a review based primarily on genetics came to another result. Atlas and Red List of the Reptiles of South Africa, Lesotho and Swaziland. Status of the Nile monitor (. The act or condition of passing winter in a torpid or resting state, typically involving the abandonment of homoiothermy in mammals. Most wild populations exhibit steady, indeterminate growth throughout their lives, but some heavily exploited populations may undergo significant growth up until 30 months of age and then cease. The diversity of these proteins in the natural world and their subsequent neofunctionalisation in venom systems presents a great opportunity to the drug discovery scientist to find structurally improved stability in a molecule by studying the venom orthologue of the CRiSP of interest. Clinical examinations for eosinophilic meningitis patients. The broad, muscular neck of the gorilla is what generates the power for their bite, but they dont use it to crush the bones of prey like the other animals we have discussed so far. Shaking movements of the head may be used to dismember prey grasped by the teeth. Nile monitors are a polygynandrous species and will mate promiscuously. What happens if a Nile monitor bites you? In 1942, Robert Mertens moved them both into the Nile monitor (V. niloticus); as synonyms or as a valid subspecies. In my spare time, you can find me getting out into nature. Two of these pig species, the endemic Bornean Sus barbatus barbatus and the only non-endemic pig species in South East Asia, the Sumatran Sus scrofa vittatus are not endangered. Aquatic, Semi-Aquatic and Riparian Vertebrates, Clinical Anatomy and Physiology of Exotic Species. Polar bears can grow up to 1,200 pounds and stand seven feet tall. What animal has the strongest bite force? Protein levels usually do not exceed 500mg/dL. 6.64), and they can reach lengths of up to 2.5cm. You canWhatsAppus on 07810 791 502. Typically, when a person suffers a minor cut, the area surround the wound will become red, swollen and warm to touch. Journal of Herpetology, 34: 414-424. Maturity is reached in three to four years. They may get defensive when attacked. Their throats and undersides are an ochre-yellow to a creamy-yellow, often with faint barring. The Nile monitor (Varanus niloticus) is one of Africas largest lizards and also one of the continents most voracious predators (for its size). These functions may vary markedly as the animal progresses from egg to juvenile to breeding adult, and may be dramatically affected by seasonal changes, especially hibernation and estivation. Tooth attachment is subpleurodont and ankylosed at an angle that minimizes stress when the teeth are loaded vertically (Rieppel, 1979). They also share over 98% of their DNA structure with modern-day humans. Figure 6.68. "There are few lizards less suited to life in captivity than the Nile monitor. American alligators can be visually identified by the fact that their lower jaw and teeth fit entirely within their upper jaw leaving no lower teeth visible when their mouths are closed. Inexperienced reptile owners often find it difficult to keep these large lizards. Dowell, S.A, D.M. Every animal has to eat, and there are few things more terrifying than staring down the maw of a hungry predator. [7], Members of the Nile monitor species group were already well known to Africans in ancient times. They have rounded nostrils which are located slightly nearer to the eye than to the snout. When the rain comes the females dig holes in the sides of termite mounds and lays their eggs. "The Sun", "Sun", "Sun Online" are registered trademarks or trade names of News Group Newspapers Limited. Contact the AZ Animals editorial team. The female lays their eggs in a pot-shaped nest built in a tree some distance away. Nambali Valsalan Vinithkumar, Nambali Valsalan Sujathkumar, in Biodiversity and Climate Change Adaptation in Tropical Islands, 2008. Nile monitors are among the most exploited species of tetrapods in Africa where it is used for food and leather. Young Nile monitors make easy prey for a large python but even larger Nile monitors are at risk. (Branch, 1998; Steel, 1996), Nile monitors inhabit a wide variety of habitats including woodland, dry savanna, scrub, evergreen thickets, swamps, and mangroves. In Indian waters five species of marine turtles recorded which include loggerhead turtle Caretta caretta (Dash and Kar,1990; Kannan and Rajagopalan,2012). The African savannah monitor (Varanus exanthematicus) is smaller than the water monitor. They are also the most dangerous large cats to humans. Being so large, you might have wondered: Are monitor lizards are dangerous to humans? Most times, monitor Lizards, will not attack unless they are provoked. She owns and operates a small farm in upstate New York which she shares with three dogs, four donkeys, one mule, and a cat. Bartlett, R., P. Bartlett. The first entry on our list is the grizzly bear (Ursus arctos horribilis). Exposed, open areas are crucial habitat components as they require sufficient basking locations. The motion of the upper jaw drives the recurved teeth into the prey. (Bartlett and Bartlett, 1996; Branch, 1998; Sprawls, et al., 2002; Steel, 1996), Nile monitors have few predators, but pythons prey on these lizards most often. ); uxamu (IsiZulu); nkwahle (Xitsonga). A cineradiographic study of feeding by V. bengalensis indicates that the upper jaw is elevated at the mesokinetic joint as the mouth opens and depressed as it closes. This species is venomous and its bite can kill its prey. The head has an elongated snout, and the tail is much longer than the body and is laterally compressed, with a low dorsal crest. [20] They have muscular bodies, strong legs, and powerful jaws. Referring to a burrowing life-style or behavior, specialized for digging or burrowing. - BBC Science Focus Nile monitors are also listed under the Endangered Species Decree of 1985, which means that international trade of the species in prohibited. Hard-shelled mollusks are a conspicuous feature of Nile monitors' diets and a thickened skull, blunt crushing teeth, and bowed lower jaw make them well-equipped for breaking open thick shells. Species groups build bowers according to one of the three basically different plans: simple platforms, avenue bowers, and maypole bowers. This species is able to grow larger than 8 feet and are known for their dangerous bite. All the species belonging to this family of Paradisaeidae show gorgeous accoutrements in form of coloured brest shields, like the Magnificent riflebird Ptiloris magnificus, elongated wiry tail feathers, like the Raggiana Bird of Pradise, Paradisaea raggiana or flank plumes, like the King Bird of Paradise Cicinnurus regius. They have a long, forked tongue like that of snakes that, together with a large Jacobsons organ, plays a major role in detecting prey by olfaction. Figure 6.63. Nile monitors are occasionally kept as pets by reptile hobbyists, but these large and aggressive lizards are a poor choice as a pet, except perhaps for very experienced keepers who can provide large enclosures. Nile monitors are occasionally kept as pets by reptile hobbyists, but these large and aggressive lizards are a poor choice as a pet except perhaps for very experienced keepers who can provide large enclosures. Varanids achieve their greatest diversity in the Northern Territory of Australia, where most of the 26 species present on the continent are found. Maybe that would have encouraged stupid dad to Monitor lizards commonly kept as pets and iguanas produce venom, according to surprising new research that is rewriting the story of lizard and snake evolution. Monitor lizards (Varanidae), originated over 90 million years ago in northern Asia (Bennett, 1995) and expanded during the Miocene to reach Europe, Africa and Australia. "They had eaten his face, eaten his hands, they actually had eaten into his abdomen and eaten some of the internal organs.". Nile monitors are also listed under the Endangered Species Decree of 1985, which means that international trade of the species in prohibited. [17][18] In Lake Chad, 336 Nile monitors had a median Ranging in weight between 3,000 to 8,000 pounds, the third-largest land mammal can grow up to 14 feet long and lives for 40 years. Saltwater crocs are responsible for 3,000-5,000 human deaths each year. On the other hand, smaller monitors are not as dangerous, but can still deliver a painful bite. In 1975, some North African countries replaced the Wild Animals Law created in 1963 with a new law that gives protection to crocodiles, pythons and monitor lizards. Dentition of a juvenile African savannah monitor (Varanus exanthematicus) with sharp pointed teeth. Pursuant to section 120.74, Florida Statutes, the Fish and Wildlife Conservation Commission has published its2022 Agency Regulatory Plan. Learn more about us & read our affiliate disclosure. Respiratory (Lower) Tract Disease/Pneumonia, Clinical Veterinary Advisor: Birds and Exotic Pets, Hormones and Reproductive Cycles in Lizards, Hormones and Reproduction of Vertebrates: Reptiles, Lenk, Eidenmueller, Staudter, Wicker, & Wink, 2005, Jacob & Ramaswami, 1976; Wikramanayake & Dryden, 1988; Phillips & Millar, 1998, infection is a history of eating uncooked fresh water snails, crustaceans, or, Maleewong et al., 2001; Eamsobhana and Yong, 2009, Utilisation of compounds from venoms in drug discovery, Cysteine rich secretory proteins (CRiSP) are predominantly found in snake venoms and some arthropods, but also in the venomous saliva of, Marine Ecosystems of Andaman and Nicobar Islands Species Abundance and Distribution, Biodiversity and Climate Change Adaptation in Tropical Islands. Their care presents particular problems on account of the lizards' enormous size and lively dispositions. Some large species like the Asian water monitor and black-throated monitor can make a unique pet for the right reptile owner. Nile monitors are carnivores with an almost insatiable appetite, they are renowned for eating just about anything they can overpower or find as carrion. Photo This may take 2-3 days to complete. You can help by taking a picture, noting the location, and reporting this information using the free IveGot1 mobile app, calling IVE-GOT1 (888-483-4681), or by reporting online at. The Florida Fish and Wildlife Conservation Commission (FWC) currently monitors local populations of breeding Nile monitors in Miami-Dade, Palm Beach and Lee Counties. In mammals these proteins play a major role in the mammalian reproductive system, such that the venom versions may have utility in reproductive medicine. Nile monitors use both visual and olfactory cues when sensing the environment. What should I do if I see a Nile monitor? Sepsis, or blood poisoning, is always triggered by an infection. An eosinophil count of more than 798 cells led to sensitivity, specificity, positive-predictive, and negative-predictive values of 76.6%, 80.2%, 58.1%, and 90.5%, respectively, for the diagnosis of meningitic angiostrongyliasis (Sawanyawisuth et al., 2010b). The lashing tail can inflict a painful wound. Herpetology, a branch of zoology that deals with the study of amphibians and reptile provide significant insight of terrestrial and aquatic ecosystem. Ivermectin-induced hepatic lipidosis in chelonians is a well-documented example. When not working with her own animals and tending her farm, Krishna is helping other animal owners with behavior or management issues and teaching neighboring farmers about Regenerative Agriculture practices. Figure 6.67. WebExotic Lizards. Nile monitors are not native to Florida and are considered an invasive species due to their impacts to native wildlife. Juvenile monitors will tail whip as a defensive measure, and as adults they are capable of inflicting moderate to serious wounds from biting and scratching. Nile monitors may be observed basking on rocks and branches and are often seen in or close to water. The Bengal or common Indian monitor (Varanus bengalensis) usually has 89 premaxillary teeth, 1011 teeth on each maxilla, and 1112 on each dentary. Clickhereto upload yours. [6] A feral population of the lizards, descended from escaped or intentionally released pets, has become established in several locations in Florida. Although they look similar to dogs in appearance, they are more closely related to cats and civets. Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes). A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. Predatory fish, herons, snakes and even adults of their own species prey on young. (Bartlett and Bartlett, 1996; Branch, 1998; Buffrenil and Castanet, 2000; Sprawls, et al., 2002; Steel, 1996), Nile monitors have been known to acclimate to human towns, farms, and homesteads, and occasionally attack livestock such as chickens. Three species [Varanus bitatawa, Varanus mabitang, and Grays monitor (Varanus olivaceus)] are arboreal and subsist largely on fruit. Monitor lizards belong to the genus of Varanus and are some of the largest lizards in the world. Barry Berkovitz, Peter Shellis, in The Teeth of Non-Mammalian Vertebrates, 2017. The olive ridley nests only on the East coast of the Andaman Islands and 12 sites have been reported along with 3 sites in Nicobar group. After hatching, the young may need to wait for rain to soften the hard nest, or reportedly the mother monitor may return at the right time and open the nest to free the hatchlings. The white blood cells and platelets form blood clots in the tissues around the cut. Police said the species is known to bite and secretes a venom that could lead to a bacterial infection, but would not likely be fatal to humans. Reptiles with acute liver disease usually present in good body condition but with sudden-onset depression and anorexia (see Liver Disease). Bacterial infection caused by a bite is the most serious concern. Bite force-wise, however, the predator remains rather potent with a maximum chomp-force of 2,980 PSI. It was suggested that venom enters the victim along grooves on the tooth surface and would penetrate deeply into the lacerations produced by the sabre-like teeth. Crocodile monitors are aggressive and quick to bite when compared with other monitors. I love learning about the various wonders nature has to offer and have been a writer for 5 years. The lashing tail can inflict a painful wound. Call: 0800 160 999, Report complaints on environmental and/or social harms and gender discrimination arising from SANBIs activities You have the animal bite down on the device and it electronically measures Harry and Meghan evicted from Frogmore Cottage by King Charles after Spare memoir, Amy Nuttall caught cheating hubby when he bought sexy lingerie, Constance Marten and lover arrested over manslaughter, Woman found murdered is missing mum who had not been seen since before Xmas, News Group Newspapers Limited in England No. The water monitor has sharp, pointed recurved teeth and eats a wide range of mainly aquatic prey, including fish, frogs and crabs, but also birds, rodents, and snakes. Nile monitors are usually solitary, with only occasional intra-species interaction outside the mating season. 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Used for food and leather Komodo Dragon: Whats the, the Fish and wildlife Conservation Commission published... Eggs in a tree some distance away they thrive around rivers ( or periodic condition changes ) will a! Up a nest when due to their impacts to native wildlife cues when sensing the environment they require basking... Wondered: are monitor lizards belong to the snout can be raised lowered. Considered an invasive species due to their impacts to native wildlife material is based upon work supported by He. Commission has published its2022 Agency Regulatory Plan and parts of Southeast Asia is used for food and leather Ursus. Grays monitor ( Varanus ) and nearly 70 species of monitor lizard vs Komodo Dragon: Whats,! '', `` Sun '', `` Sun '', `` Sun '', `` Sun,... Monitor lizards are capable of competing with the study of amphibians and reptile provide significant insight of terrestrial aquatic... Greatest threat species prey on young terrestrial and aquatic ecosystem habitat components as thrive... And anorexia ( see liver disease ) and even adults of their own species on., reptiles, fishes, arthropods, mollusks, eggs, birds and. Related to cats and civets can be raised or lowered relative to its size the... The sea and reach sandy coastal areas to lay their eggs in the sides of mounds... Affiliate disclosure 20 ] they have rounded nostrils which are located slightly nearer the! Learning about the most serious concern Silverton, 0184 paradise ( Fam,! Grow larger than 8 feet and are known for their dangerous bite Branch, W.R.,,. Caretta ( Dash and Kar,1990 ; Kannan and Rajagopalan,2012 ) lowered relative to the of! Exanthematicus ) is smaller than the water monitor ( V. indicus is more widespread occurring in New and!, or blood poisoning nile monitor bite force is always triggered by an infection if I a. '' ( On-line ), and mollusks referring to a burrowing life-style or behavior, specialized for digging burrowing...

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